Forgiveness Synthesis

When I think of forgiveness, first thing that comes to mind is hurt.  In the human experience we have created a demand for reconciliation.  What does forgiveness look like in nature?  In the great chain of life there are processes that aren't fair, but are there processes that are expressed with malice?  I think that the nature of beings is to oscillate between balance and other vertices, always striving for balance.  Is one of those vertices malicious?  There is a sophisticated expression of injustice when it comes to people because we can reason, rationalize, or empathize.  We also struggle with the privilege of technology and I believe this makes most human injustice irrational.  Other species do most things for the sake of nutrition, order, and reproduction.  All may be narrowed to proliferation.  We are not different except for qualities like revenge, grudge, or forgiveness.  How is forgiveness expressed in nature?

What can it teach me?